ApartAdemais fromdas thedúas two living [[family`familia (biologybioloxía)|familiesfamilias]] vivas, therehai are a few prehistoricalgúns [[taxataxon]]s inprehistóricos thede Megaloptera, onlycoñecidos knownpolos fromseus [[fossilfósil]]ses. SomeAlgúns ofdestes theseocupan occupyunha aposición moremáis [[basal (evolutionevolución)|basal]] position:<ref>SeeVer referencesreferencias inen Haaramo (2008)</ref>
* GenusXénero ''[[Corydasialis]]'' (sometimesás consideredveces considerado a familia [[monotypicmonotípico|monotípica]] family [[Corydasialidae]]<ref>Engel & Grimaldi (2007)</ref>)
* FamilyFamilia [[Nanosialidae]]<ref>Shcherbakov DE (2013) Permian ancestors of Hymenoptera and Raphidioptera. ZooKeys 358: 45–67. doi: 10.3897/zookeys.358.6289</ref>
The Megaloptera are monophyletic and are a sister clade of the [[Neuroptera]].<ref name=Wang2012>Wang Y, Liu X, Winterton SL, Yang D (2012) The first mitochondrial genome for the fishfly subfamily Chauliodinae and implications for the higher phylogeny of Megaloptera. PLoS One 7(10):e47302. doi: 10.1371/journal.pone.0047302</ref> Within the Megaloptera, Corydalinae and [[Chauliodinae]] are sister clades. The divergence time estimation suggests that the stem lineage of Neuropterida and [[Coleoptera]] separated in the Early [[Permian]]
